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-Root hair length and density
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-root cortical aerenchyma
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-root tissue density (related to secondary wall development and tissue proportions)
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-specific root length
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-carboxylate exudates
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-resource allocation among root classes
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-phenology of root formation
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-root cortical senescence
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-seminal root whorls- number of seminal roots
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-shoot born rooting- both mesocotyl and stem
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-root gravitropic growth angles
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-lateral branching
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-plasticity of the above in response to nutrient and drought stress
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